威利斯假说是Dr. Willis在1915-1916年提出来的, 原本构建在植物上的假说。
大意就是：在特定的时间地点，没有明显障碍的理想情况下，越古老的物种越强大 （占据的区域更大）。
“The area occupied by any given species (of plants) at any given time in any given country in which there occur no well-marked barriers depends upon the age of that species in that country.”
明显是反达尔文的假说，这些年来也不红。
相关论文戳这里http://www.jstor.org/stable/1643607?seq=1
原始文献在这里：
1. Willis, J. C.: The Endemic Flora of Ceylon, with reference to Geographical Distribution and Evolution in general. Phil. Trans., B, vol. ccvi, 1915, p. 307. ［［on p. 209］］
2. Ibid.: The Evolution of Species in Ceylon, with reference to the Dying Out of Species. Ann. of Bot., vol. xxx, 1916, p. 1. ［［on p. 209］］
3. Ibid.: The Distribution of Species in New Zealand. Ann. of Bot., xxx, 1916, p. 437. ［［on p. 209］］
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